Birth-date: July 15, 1925

Place of birth: Lawton, Oklahoma

Education:

Lawton High School, 1942
Cameron State Agriculture College, Lawton, OK, AS., 1944
University of Oklahoma, Norman, OK, B.A. (Zoology), 1948
University of Wisconsin, Madison WI, Ph.D. (Genetics and Zoology), 1954 

Professional Experience:

• 1980- 1994 Professor of Genetics, Iowa State University
• 1978-1979 Collaborating Professor, MS-6, Universidade Estadual de Sao Paulo, Jaboticabal, Brasil
• 1962-1980 Associate Professor, Iowa State University
• 1955-1962 Associate Specialist, University of California at Davis, Department of Microbiology, School of Veterinary Medicine
• 1953-1955 Project Associate, University of Wisconsin, Department of Genetics
• 1948-1953 Research Assistantship, University of Wisconsin, Department of Genetics
• 1992-1993 Visiting Foreign research Professor, Universidade Federal de Minas Gerais, Departmento Zootecnia de Escola Verterinaria, Belo Horizonte, M.G., Brasil
• 1995-1997 Visiting Foreign research Professor, Universidade Federal de MinasGerais, Departmento Zootecnia de Escola Verterinaria, Belo Horizonte, M.G., Brasil

WJM- Wilmer J. Miller Research Discoveries;   Supplement to CV
Academic Research Origin:  M. R. Irwin’s Laboratory in Classical  Immunogenetics; University of Wisconsin, Madison.

1.      Established that antibodies are not mutagens (contrary to Guyer’s proposal), using anti-lens in rabbits. 

2.      Showed that development of species-specific antigens may have a “gap”, e.g. AF of C. guinea.

3.      Showed a displacement or replacement and non-simultaneous development of adult antigenic factors in cattle (with Hubbert).

4.      Developed “Miller’s principle of antithetical antigens”: A-A’, B-B’,C-C’, E-E’..

5.      Taxonomic implications for species-specific antigenic relationships reveal that they may be identical, related, or unique (with Irwin), and may exist as phenogroups.  

6.      Explained the “hybrid substance” as an interaction product dependent on interacting non-allelic loci heterozygous and homozygous in a species, e.g. CC’, hu=4//hu-8. 

7.      Developed the cloacal method for sexing doves and pigeons—30 species.

8.      First to list behavior traits in the ringneck dove. 

9.      Demonstrated the hormonal control of sexual dichromatism in the dwarf turtle dove, S. humilis. 

10.  Performed genetic analysis of 55 genetic characters in Streptopelia risoria, ringneck dove, and discovered mutants: 

•  heel walker                   pigeon-  recessive 

• flare                              dove   -  recessive, shows in juvenile down also

• stubby dwarf    dove   -  recessive

• silky plumage    dove   -  partial or codominant(gene  symbol L)

• thin bill ring       dove  -   partial or codominant 

• yellow down     dove  -  recessive 

Assisted in the analysis of pigeon mutants: (with Hollander) 

• amputated toes – recessive

• fray plumage – partial dominant

• sideburns – partial dominant

• web-lethal --  sex-linked recessive

11.  Conducted dihybrid testcrosses for linkage of several characters including 4 species-specific antigens and albumin of S. risoria     and S. humilis.  There  were two linkage groups found:  _L__8__hu-8_  :  and alb-H___21__ hu-y

12.  Found heterosis in antibody production- Pheasant-Turkey hybrids (with Wolfe). 

13.       Found bacterially mediated false-positive reactions in blood typing tests in vitro (with Stormont), and an in vivo influence by Salmonella in red cells of pigeons.  

14.   Assisted (with Draper) in the analysis of spinal dysraphism in dogs, which is a dominant with reduced penetrance and variable expressivity. 

14.       Dogs have blood type phenogroups; and a dog reagent with human blood types showed type AB have mandatory pairing on bifurcated chains (with Steiler). 

15.       Made rather extensive cross species comparisons showing that naturally occurring antibodies are highly frequent in cross-species tests, and that natural cattle U’ reagent (with Stone) is a U1U2 subtype heterozygote     detector. 

17.  Showed that cattle have a neonatal hemoglobin; that fetal cattle have 4 fetoproteins; that fetal cattle have mostly non-IgG antibodies, since 2 ME inactivated them (with Hubbert). 

18.    Started turkey blood typing. 

19.    Demonstrated that recessive antigens are more widespread than generally believed (with Fischer). 

20.    Was among the first to apply gel electrophoretic character–polymorphism to wild species, e.g. the white-tailed deer, using hemoglobin and transferrin differences (with Haugen & Roslien). 

21.    First to list the criteria for blood tying reagents. 

22.    Increased the B phenogroups of cattle from 120 to 300+.

23.    Showed that buffalo – bison (typed with cattle reagents) may show some of the cattle individual differences as species-specific antigens; L, including    dual alternative specificities (heterozygotes) R’S’, as a species-specific antigen (homozygotes)! 

24.    Demonstrated (with Stormont) that bison show individual and species-specific antigens in the same system, breaking the then current dogma!  specifically:                                 

                                                    C1WX₁    

                                WX₁

                                    X₁

25.    Characterized blood types (especially phenogroups) in several breeds of cattle: Longhorn, Caracu, Gir, Nelore, Guzerat, and further developed that in Aryshire.  Brown Swiss, Jersey, Guernsey, Holstein-Friesian, Aberdeen Angus, Hereford, and Shorthorn.

26.    Found new systems N and R’-S’, and several new factors in cattle blood typing. 

27.    Showed that subtypes are asymmetrical blood typing reactions widely and extensively present.  They may be

• Linear, 

• Non-linear,  

• Quantitative and Qualitative. 

28.    Screened guanacos for suitability in blood typing and for taxonomic relationships. 

29.  Performed extensive lectin tests among and within species finding similarities and differences with antibodies.  Examples:

• The peanut extract gives a recessive reaction in ringneck doves! 

• Maackia is antithetical to peanut in ringneck doves.

• Wisteria extract detects a juvenile character and shows a cluster effect in ringnecks!

• Lectins may be fractionated by absorptions, breaking the then current dogma.

• White bauhinia seed’s lectin shows individual differences in dogs. 

Membership in Professional Societies:

• AAAS (Fellow)
• American Genetics Association
• American Institute of Biological Sciences
• American Society of Naturalists
• Genetics Society of America
• New York Academy of Science
• Society of Experimental Biology and Medicine
• International Society for Animal Blood Group Research
• The American Association of Immunologists 

Membership in Honorary Societies:

Phi Sigma
Sigma Xi
Phi Theta Kappa
Gamma Sigma Delta 

Refereed Publications-

1. Miller, W. J. 1953(a). Immunogenetic studies of cellular antigens of species and species hybrids in Columbidae. Ph.D. Thesis, University of Wisconsin. 96 pp.
2. Miller, W. J. 1953(b). The time of appearance of species-specific antigens of Columba guinea in the embryos of back-cross hybrids. Physiological Zool. 26: pp. 124- 131.
3. W. J. Miller and Bryan, C. R. . 1953(a). Serological differentiation of the homozygotes and heterozygotes in back-cross birds following a species cross in Columbidae. Proc. Nat. Acad. Sci. 39: pp. 407-412.
4. Bryan, C. R. and W. J. Miller. 1953(b). Interaction between alleles affecting cellular antigens following a species cross in Columbidae. Proc. Nat. Acad. Sci. 39: pp. 412- 416.
5. Miller, W. J. and F. H. Wagner. 1955. Sexing mature Columbiformes by cloacal characters. The Auk 72: pp. 279- 285.
6. Miller, W. J. and W. H. Stone. 1955. Blood typing the erythrocytes of bison with reagents for antigenic factors of cattle. Genetics 40: p. 586.
7. Miller, W. J. 1956. Silky plumage in the ring neck dove. Jour. Hered. 47: pp. 37-40.
8. Miller, W. J. 1956. The hybrid substance of the erythrocytes of the hybrids between Columba livia and Streptopelia risoria. Genetics 41: pp. 700-714. 
9. Miller, W. J. 1958. Anti-lens sera as a mutagen in rabbits. Jour. Exper. Zool. 137: pp. 463-477.
10. Miller, W. J., and L. S. Miller. 1958. Synopsis of behaviour traits of the ring neck dove. Animal Behaviour 6: pp. 3-8.
11. Miller, W. J. 1960. Sexual dichromatism in two species of the Columbidae. The Condor 62: pp. 45-48.
12. Gregory, P. W. and W. J. Miller. 1960. Coat color, pattern horn condition and blood groups in cattle. Genetics 45: p. 1000.
13. Miller, W. J., C. Stormont, and Y. Suzuki. 1960. Bacterially mediated false-positive reactions of lytic blood-typing tests. Nature 186: pp. 247-248.
14. Miller, W. J. 1961. A second egg tooth in a mourning dove. The Auk 78: p. 275.
15. Irwin, M. R. and W. J. Miller. 1961. Interrelationships and evolutionary patterns of cellular antigens in Columbidae. Evol. 15 (1): p. 3043.
16. Miller, W. J., and W. H. Stone. 1961. Naturally occurring isoantibodies of the S blood group system in cattle. J. Immunol. 86: pp. 165-169.
17. Miller, W. J., C. Stormont, and Y. Suzuki. 1961a. Blood groups and the taxonomic status of American buffalo and domestic cattle. Evol. 15: pp. 196-208.
18. Miller, W. J., C. Stormont, and Y. Suzuki. 1961b. The S system of bovine blood groups. Genetics 46: pp. 541-551.
19. Asmundson, S., W. J. Miller, and H. R. Wolfe. 1961. Antibody production in pheasant- turkey hybrids. Poultry Sci. 40 (1): pp. 140-144.
20. Miller, W. J. 1964. First linkage of a species antigen in the genus Streptopelia. Science 143: pp. 1179-1180.
21. Miller, W. J. 1965. A recessive lectin-factor in Streptopelia risoria. Genetics 51: pp. 247-251.
22. Haugen, A. O., W. J. Miller, and D. J. Roslien. 1965. Natural variation in the blood proteins of white-tailed deer. Jour. of Wildlife Management 29: pp. 717-723.
23. Hollander, W. F. and W. J. Miller. 1965. Homology of color mutants in two species of ducks. Jour. of Heredity 56: pp. 295-297.
24. Asmundson, V. S., G. R. J. Law, W. J. Miller and C. Stormont. 1965. Blood groups of turkeys. Genetics 51: pp. 151-158.
25. Miller, W. J. 1966. Evidence for two new systems of blood groups in cattle. Genetics 54: pp. 151-158.
26. Miller, W. J. 1966. Blood groups in longhorn cattle. Genetics 54: pp. 391-404.
27. Miller, W. J. 1966. Hemagglutinins in normal sera and species relationships. Iowa State Journ. of Science 41: pp. 181-190.
28. Miller, W. J. 1967. Segregation of plasma-albumin types from a species cross. Proc. Soc. Exp. Biol. & Med. 124: pp. 1224-1227.
29. Miller, W. J., and I. R. Quinteros. 1968. An alternative method in distinguishing cattle transferrin phenotypes. Biochem. Genetics 2: pp. 213-218.
30. Miller, W. J., and I. R. Quinteros. 1968. Inhibicion de algunos "reactivas especificos" de grupos sanguineas bovines, mediante diferentes azucares. Rev. Fac. C. Vet. La Plata Anox - No. 23 – III2 Epoca - May-Agosto 191: p. 200.
31. Miller, W. J., and J. L. Weber. 1969. First case of non-allelic interaction product as a species-specific red cell antigen. Genetics. 62: pp. 619-623.
32. Miller, W. J. and I. R. Quinteros. 1969. Busqueda de antigenos holandricos o ligados al chromosoma "Y". Analecta Veterinaria No. 2, 1: pp. 59-67.
33. Hubbert, W. T. and W. J. Miller. 1971. Development polymorphism in bovine hemoglobin. Amer. Jour. Vet. Res. 32: pp. 1723-1730.
34. Hubbert, W. T. and W. J. Miller. 1972(a). Naturally occurring antibody in bovine fetal serum: reactivity against homologous and heterologous species erythrocytes.
35. Jour. Animal Blood Groups and Biochemical Genetics 3: pp. 3-17.
36. Hubbert, W. T. and W. J. Miller. 1972(b). Alpha-Fetospecific serum proteins in bovine fetuses. Proc. Soc. Exp. Biol. & Med. 141: pp. 685-687.
37. Hubbert, W. T. and W. J. Miller. 1974. Immunogenetic ontogeny of cellular membrane function: A review. J. Cell Physiol. 84: pp. 429-444.
38. Hubbert, W. T. and W. J. Miller. 1975. Adult isoantigen and lectin reactivity of bovine fetal red cells. Biology of the Neonate 27: pp. 23-39.
39. Hollander, W. F. and W. J. Miller. 1975. The ABO’s of blood types. Iowa Sci. Teachers Jour. 12: p. 24.
40. Draper, D. D., J. P. Kluge, and W. J. Miller. 1975. Neurologic, pathologic and genetic aspects of spinal dysraphism in dogs. Zbl Vet. Med., Reihe C. Bd. 4, Heft 4.
41. Miller, W. J. 1976. Blood groups: Why do they exist? BioScience 26: pp. 557-562.
42. Miller, W. J. 1977. Purebred!?. Iowa Science Teachers Journal 14: 37-38. Purebred!?. ADAN Sept./Oct. 1991. Reprinted ADAN Nov./Dec. 1998.
43. Miller, W. J., R. J. Moreno, I. R. Quinteros, and E. D. Tejedor. 1977. Comparative racial study of blood genetic markers in the Argentine creole cattle. Abs.III
44. Latin American Congress of Genetics, Montevideo, Uruguay, 2-5 Feb. and 6-12 Feb. Eds. M. E. Drets, N. Brum-Zorilla, and G. A. Folle NSF-UNESCO, OAS.
45. Hollander, W. F. and W. J. Miller. 1978. Amputated, frayed, and sideburns; three new mutants in the pigeon. Iowa State Journal of Research Nov. Vol. 53. No. 2: pp. 109-118.
46. Hollander, W. F. and W. J. Miller. 1981. Hereditary variants of behavior and vision in the pigeon. Iowa State Journal of Research 55: pp. 232-331.
47. Miller, W. J. 1983. Pedigree style in teaching genetics. Proc. Iowa Acad. Sci. 90 (1): pp. 26-27.
48. Bortolozzi, J., H. C. Hines, W. J. Miller, I. R. Quinteros, M. A. Poli, F. Sal Paz. and D. E. Tejedor. 1983. Segregation of transferrins in Creole cattle in Argentina. Indian Vet. Jour. Feb. 60: pp. 93-98.
49. Assis, Ana Isabel, Miriam Luz Giannoni, W. J. Miller, and F. L. Pires. 1984. Avaliação dos tipos sanguineos do Caracu, o gado crioulo Brasileiro. B. Indúst. anim., Nova Odessa, SP 41 (Único): 1-8.
50. Miller, W. J. 1985. Appropriate gene symbols in teaching genetics. Proc. Iowa Acad. Sci. 92 (3): pp. 115-118.
51. Franklin, W. J., P. J. Hollander, and W. J. Miller. 1985. Blood typing South American camelids. J. Heredity. 76 (5): pp. 369-371.
52. de Carillo, I. P., P. J. Hollander, and W. J. Miller. 1986. Further search for interaction antigens on erythrocytes in selected species hybrids of birds. Poultry Science 65: pp. 1028-1033.
53. Ramirez, German, MD ; Wilmer. J. Miller, PhD;, Polly A. Bittle, BSN; Alvaro Hidalgo,
54. ??DVM; Rafael Santacruz, DVM; Gene Colice, MD. Blood types in cattle of Iberian ancestry and in Holsteins at various altitudes. Am J Vet Res 53 No. 7 July : p.1248-1252.
55. Hollander, Willard F., and W. J. Miller. 1995. Three neglected advances in classical genetics. BioScience: Feb. 1995. Vol. 45, No. 2.
56. Miller, W. J. 1996. Dominance. Brazilian Journal of Genetics: 19 (4): p. 677.
57. Miller, W. J. 1997. Dominance, codominance and epistasis. Brazilian Journal of Genetics: 20 (4): pp. 663-665. 
58. Miller, Wilmer J. 1997. O Pombo Doméstico Feral, Columba livia, um estudo de genetica. Actualidades Ornitologicas-Set/Oct-1997.

 Popular Articles-

1. Miller, W. J. 1969. The biology and natural history of the mourning dove. Public Forum address sponsored by Ames Audubon Soc.
2. Hollander, W. F., and W. J. Miller. 1975. The ABO's of blood types. Iowa Science Teachers Journal 12:24 (No. 4 Dec.).
3. Miller, W. J. 1980. A visit with a pigeon breeder in Brazil. Invited talk at the American Pigeon Fanciers Council. 25 July 1980. St. Louis, Missouri.
4. Hollander, W. F. and W. J. Miller. 1982. A new sex-linked mutation "web-lethal" from racing homers. American Racing Pigeon News Oct. pp. 50-51.
5. Miller, W. J. 1984. Basic color mutants of the ring neck dove and data on their inheritance under the title "Doves and Such". 7 April 1984. Invited talk at the Wisconsin Pigeon Breeder's Council.

ADAN Articles (American Dove Association Newsletter)-

1. Miller, W. J. 1974. A recessive mutant "stubby dwarf" in ringneck doves. ADAN Sept./Oct. pp. 5-8.
2. Miller, W. J. 1976. How heavy are your doves? ADAN Sept./Oct. p. 7.
3. Miller, W. J. 1977. White head flecks in doves. ADAN Jan./Feb. p. 1. Reprinted Jan./Feb. 1981.
4. Miller, W. J. 1978. Absence of bill ring in ringneck doves, Streptopelia risoria. ADAN May/June p. 4. Reprinted ADAN July/Aug. 1980 p. 6.
5. Hollander, W. F. and W. J. Miller. 1979. New colors in Laughing-doves. ADAN May/June p. 4.
6. Hollander, W. F. and W. J. Miller. 1979. The story of the pied mutations. ADAN Jan./Feb. pp. 8-9.
7. Miller, W. J. 1979. Breeding cycle interval in ringneck doves, Streptopelia risoria.
8. ADAN March/April p. 7.
9. Miller, W. J. 1980. A further experience with disease in doves. ADAN Jan./Feb. p. 4.
10. Miller, W. J. 1980. Dove Culture V. What do they eat?. ADAN Jan./Feb. pp. 9-11.
11. Miller, W. J. 1980. Fostering dove eggs. ADAN May/June p.3.
12. Hollander, P. J. and W. J. Miller. 1981. Do some of your ring neck doves have long tails? ADAN May/June p. 5.
13. Hollander, P. J. and W. J. Miller. 1981. Down color in ringneck doves, Streptopelia risoria. ADAN July/August pp. 4-5.
14. Miller, W. J. 1981. Cream colored ringnecks. ADAN Nov./Dec. p. 6.
15. Miller, W. J. 1981. Termination of dove colony and genetic characters studied. ADAN Nov./Dec. pp. 7-8.
16. Miller, W. J. 1984. A record holder per number of progeny!? ADAN March/April pp. 8-9.
17. Miller, W. J. 1984. Book review of "Wild Pigeons and Doves" by Dr. Jean Delacour
18. 1980. T.F.H. Publications. pp. 189. ADAN March/April p. 7.
19. Miller, W. J. 1984. Genetics of the ringneck dove, Streptopelia risoria I. Overveiw. ADAN May/June p. 10.
20. Miller, W. J. 1984. Genetics of the ringneck dove, Streptopelia risoria II. Description of mutants- albino, autosomal recessive= al. ADAN Sept./Oct. pp. 3-5.
21. Miller, W. J. 1984. Genetics of the ringneck dove, Streptopelia risoria III. Description of mutants- Ivory- autosomal recessive= iv. ADAN Nov./Dec. pp. 6-8.
22. Miller W. J. 1985. Genetics of the ringneck dove, Streptopelia risoria. IV. Description of mutants- pied- autosomal recessive. ADAN Jan./Feb. pp. 9-11. Reprinted ADAN May/June 1988 pp. 5-6.
23. Miller, W. J. 1985. Sleeping doves. ADAN Nov./Dec. p. 14. Reprinted ADAN Nov./Dec. p. 8.
24. Miller, W. J. 1985. Cream: the first ivory-rosy ringneck doves. ADAN Sept./Oct. pp. 6-8.
25. Miller, W. J. and O. F. Munsell. 1985. Questions and characteristics of orange- tangerine ringneck doves. ADAN Nov./Dec. p. 5. Reprinted ADAN May/June 1993 p. 6.
26. Miller, W. J. 1986. Cold-weather blood feathers in doves?. ADAN Jan./Feb. p. 10.
27. Miller, W. J. 1986. The wild type (dark) color. ADAN Jan./Feb. p. 6.
28. Hollander, W. F. and W. J. Miller. 1986. Silky revisited. Pigeon Genetics: news, views, documents, issue 13 March 1986 pp. 3-4.
29. Miller, W. J. 1986. The fate of the world depends on the collared dove. ADAN March/April p. 7.
30. Miller, W. J. 1986. Early domestic Doves in the New World. ADAN Sept./Oct. p. 10. Reprinted ADAN Nov./Dec. 1986 p. 10.
31. Miller, W. J. 1987. Comments on the new mutant colors in Ringnecks. ADAN Jan./Feb. p. 8.
32. Miller, W. J. 1987. Cream and Albino Doves. ADAN May/April p. 4. Cream and Albino Doves. The Doveline Sept./Oct. 1998 p. 5.
33. Miller, W. J. 1987. A note about minerals and diet for doves. ADAN Nov./Dec. p. 5. Reprinted ADAN May/June 1992 pp. 2-3.
34. Hochlan, Gene, and W. J. Miller. 1988. Same sexed clutches in mourning doves?. ADAN May/April pp. 10-11.
35. Miller, W. J. 1989. Dark-Blond-White: Sex-linked alternatives in ringneck doves. ADAN Jan./Feb. pp. 6-10.
36. Hollander, W. F. and W. J. Miller. 1989. Mutation in ringneck doves. ADAN March/April pp. 7-8.
37. Miller, W. J. 1989. New mutant color in ringneck doves. ADAN Sept./Oct. pp. 3-4.
38. Miller, W. J. 1990. A possible new mutant, hallux nail spike, in ringneck doves. ADAN July/Aug. p. 5.
39. Miller, W. J. and K. L. Stalder. 1990. Pink Ringneck doves!!! ADAN Sept./Oct. pg.10. Pink Ringneck doves. The Doveline Sept./Oct. 1998 p. 5.
40. Miller, W. J. 1990. About frosty color and how to tell frosty at hatching. ADAN Nov./Dec. 1992
41. Miller, W. J. 1991. Long term production in a white dove pair, Streptopelia risoria. ADAN Sept/Oct. pp. 5-6.
42. Miller, W. J. 1991. Long term production in a rosy pair, Streptopelia risoria. ADAN Nov./Dec. pp. 10-11.
43. Miller, W. J. 1992. Distinguishing (?) the cream phenotype in doves. ADAN Jan./Feb. pp. 7-8.
44. Miller, W. J. 1992. Rosy ringneck doves and other color comments. ADAN May/June p. 4.
45. Hollander, W. F. and W. J. Miller. 1996. Book review of "Doves: A Complete Pet Owner’s Manual" by Matthew M. Vriends 1994. Published by Barron’s pp. 115. ADAN May/June p.12.

PS & GN Articles (Pigeon Science and Genetics Newsletter)-

1. Miller, W. J. 1976. Ivory = opal? PS & GN 2:24.
2. Miller, W. J. 1976. Appropriate gene symbols. PS & GN 2:18.
3. Miller, W. J. 1976. Sexing doves III. Sex-linked genetic characters. PS & GN 2: 32-33.
4. Miller, W. J. 1976. Sexing doves IV. Other methods. PS & GN 3: 11-12.
5. Miller, W. J. 1976. Reproductive success (?) in pigeons. PS & GN 3:26-29.
6. Hollander, W. F. and W. J. Miller. 1977. Symbols in Genetics. PS & GN 5: 47
7. Miller, W. J. 1977. A helpful treatment for salmonella in doves. PS & GN 6: 18.
8. Miller, W. J. 1978. Dove culture I. Incubating eggs. PS & GN 7: 24-26.
9. Hollander, W. F. and W. J. Miller. 1978. The quest for linkages. PS & GN 8: 13-14.

Articles published in ADAN and PS & GN-

1. Miller, W. J. 1974. Turn on the lights?. ADAN Nov./Dec. pp. 5-7. Turn on the lights? PS & GN 1: 26-28. Reprinted ADAN Sept./Oct. 1977 pp. 5-6.
2. Miller, W. J. 1974. More about dove diseases. ADAN Nov./Dec. pp. 3-4. More about dove diseases. PS & GN 1: 36-37.
3. Miller, W. J. 1974. Sexing doves I. Vent sexing. ADAN Sept./Oct. 1974 p. 3. Sexing doves I. Vent sexing. PS & GN 1:20. Sexing doves I. Vent sexing. The Doveline Nov./Dec. 1998 p. 12.
4. Miller, W. J. 1977. Dove culture II: Differences in baby ringneck doves. ADAN July/Aug. p. 5. PS & GN 1978 8:7.
5. Miller, W. J. 1977. Dove culture III: Differences in juvenile ringneck doves. ADAN July/Aug. p. 6. PS & GN 1978 8: 8.
6. Miller, W. J. 1977. Plication (tailfolding) in ringneck doves. ADAN Nov./Dec. p.5. PS & GN 7: 39.

Articles in ADAN, PS & GN and American Pigeon Journal-

1. Miller, W. J. 1974. Sexing doves II. Sexing by behavior. ADAN July/Aug. p. 3. Reprinted ADAN May/June 1992 and Jan./Feb. 1996. PS & GN 1: 21. 1978. American Pigeon Journal 67: 139. The Doveline Jan./Feb. 1999. p. 12. 

American Pigeon Journal Articles-

1. Hollander, W. F. and W. J. Miller. 1967. Mutations in ringneck doves. American Pigeon Journal 56: 135.
2. Miller, W. J. 1974. Body Temperatures of adult pigeons. American Pigeon Journal July: 397.
3. Miller, W. J. 1981. A visit with a pigeon fancier in Brazil. American Pigeon Journal April 1981. 29-30.

Articles in ADAN and American Pigeon Journal-

1. Miller, W.J. 1974. Predation on doves. ADAN July/Aug. pp. 6-9. Watch for predators on doves. American Pigeon Journal Feb. 64: 111.

PGNL Articles (Pigeon Genetics Newsletter)-

1. Miller, W. J. 1969. First linkage of a species antigen in genus Streptopelia. PGNL No. 50 pp. 30-31.
2. Miller, W. J. 1969. The biology and natural history of the mourning dove. PGNL No. 50: pp. 50-51.
3. Miller, W. J. 1971. Allelism of pigeon and dove albinism plus interaction of rosy and pied with white in doves. PGNL No. 59, p. 2-3.
4. Miller, W. J. 1972. Test of allelism and recessive red of pigeons and rosy in doves. PGNL No. 62, p. 33.

 Abstracts-

1. Bryan, C. R. and W. J. Miller. 1951. Interactions between alleles to produce a hybrid substance. Genetics 36: 544-545. Abstract.
2. Miller, W. J. and W. H. Stone. 1953. A new normal antibody of cattle serum. Genetics 38: 693. Abstract.
3. Miller, W. J. 1954. Segregation of species-specific antigens and the hybrid-substance in back-cross hybrids following a genetic cross in Columbidae. Genetics 39: 983- 984. Abstract.
4. Miller, W. J. and W. H. Stone. 1955. Alterations in the reactivity of cattle erythrocytes following treatment with enzymes. Genetics 40: 599. Abstract.
5. Miller, W. J. and C. Stormont. 1957. Parentage exclusions by blood grouping in dairy cattle. Genetics 42: 385-386. Abstract.
6. Miller, W. J., C. Stormont and Y. Suzuki. 1957. New phenogroups in the B system of bovine blood groups. Genetics 42: 400. Abstract.
7. Miller, W. J. 1960. Blood groups in longhorn cattle. Genetics 45: 1000. Abstract.
8. Law, G. R. and W. J. Miller. 1960. The A system of turkey blood groups. Genetics 45: 997. Abstract.
9. Asmundson, V. S., G. R. Law and W. J. Miller. 1960. Blood types of turkeys. Poultry Science 39: 1268. Abstract.
10. Miller, W. J., C. Stormont and Y. Suzuki. 1960. The convergence of the A-H and D systems of bovine blood groups. Genetics 45: 1013. Abstract.
11. Miller, W. J. 1961. Evidence for two new systems of blood groups in cattle. Genetics 46: 883. Abstract.
12. Ham, W. E., W. J. Miller and C. C. O' Mary. 1961. Further studies on albinism in beef cattle. Amer. Soc. of Animal Production Western Meetings, November 24. Abstract.
13. Miller, W. J. 1962. Hemagglutinins in normal sera and species relationships. Collected reports from the 8th Animal Blood group Conference in Europe, Ljubljana, Yugoslavia, Aug. 21-24. pg. 8.
14. Miller, W. J. 1963. The agglutination of erythrocytes of ringneck doves with peanut extract. Genetics 48: 901-902. Abstract.
15. McKeever, S. and W. J. Miller. 1964. Lectin-deteced differences among wild rodents. Genetics 50: 269. Abstract.
16. Draper, Donald D., W. J. Miller and Mary Shelton. 1976. Genetics aspects of spinal dysraphism. Presented at the Stadler Symposium, University of Missouri.
17. Hollander, W. F. and W. J. Miller. 1977. Observations on pigeons having congenital head tremor associated with the "Sideburns" mutant. Behaviour Genetics Assoc. 7th Annual Meeting, 27-30 April. Abstract.
18. Miller, W. J., I. R. Quinteros, and E. D. Tejedor. 1977. Fish transferrins of the order Siluriformes. Abs. III Latin-Amer. Cong. Gen.; Montevideo, Uruguay, 2-12 Feb.
19. Larramendy, R. H., W. J. Miller, I. R. Quinteros, and E. D. Tejedor. 1977. Some rare transferrins in bovines. Abs. III Latin-Amer. Cong. Gen.; Montevideo, Uruguay, 2-12 Feb.
20. Huca, G. A., Miller, W. J., I. R. Quinteros, F. Sal Paz, and E. D. Tejedor. 1977. Segregation of blood phenogroups in the Argentine "Bovine criollos". Abs. III. Latin-Amer. Cong. Gen.; Montevideo, Uruguay, 2-5 and 6-12 Feb.
21. Miller, W. J. 1978. Legions tipos silvestres y consideraciones geneticas del longhorn norteamericano. Abs. VI Jornadas Internacionales de la Facultad de Ciencias Veterinarias de la Universidad Nacional de La Plata 5-11 Nov. 78.
22. Miller, W. J., I. R. Quinteros, and E. D. Tejedor. 1978. Investigaciones Immungeneticas en el bovino criolo Argentino-Marcadores geneticos. Abs. VI Jornadas Internacionales de la Facultad de Ciencias Veterinarias de la Universidad Nacional de La Plata 5-11 Nov. 78. 
23. Miller, W. J. and Carol M. Stieler. 1979. Evidence from lectin reactions for mandatory pairing of the terminal sugars in human blood group AB. Posterboard presentation, Sociedade Brasileira de Bioquimica VIII Reuniao Annual 18-21 April 1979, Caxambu, M.G. Brasil.
24. Carillo, I., P. J. Hollander and W. J. Miller. 1984. Tests for interaction product in chicken/Guinea fowl hybrids. I.A.S. meetings, Iowa City, Iowa, 22 April 84.
25. Miller, W. J. and K. A. Stille. 1984. Cross-reactive patterns among cattle and goat erythrocytes in antigen activity. I.A.S. meetings, Iowa City, Iowa, 22 April 84.
26. Graupera, K. S. and W. J. Miller. 1984. Reactivity of antisera from rabbits against goat erythrocytes. I.A.S. meetings, Iowa City, Iowa, 22 April 84.
27. Miller, W. J., T. A. Olson and K. Stille. 1985. Blood types of Florida Scrub cattle and their comparison with other Iberian source stocks. G.S.A. meeting, Boston, Massachusetts Genetics 110: Supplement No. 3, Pt. 2, s80.
28. Miller, W. J. and K. Stille. 1985. Production of immune reagents for blood typing goats. I.A.S. meetings, Wartburg, Iowa, 97th Session, 27-28 April 1984.
29. Miller, W. J. and W. Schauer. 1985. Normal antibodies in blood typing goats, sheep and cattle. I.A.S. meetings, 97th Session, 27-28 April 84. 
30. Miller, W. J. and K. A. Stille. 1986. Immunogenetic relationships of erythrocytic antigens of goats and cattle. G.S.A. meetings, Urbana, Illinois, 55th Session, 8-10 June, 1986.
31. Miller, W. J. 1986. Mendelian segregation and assortment for 7 genetic characters in ringneck doves, Streptopelia risoria. G.S.A. meetings, Urbana, Illinois, 55th Session, 8-10 June, 1986.

Newsletter Articles-

1. Miller, W. J. 1959. Antigenic factors and blood grouping techniques. The Immunogenetics Letter Vol. 1, No. 2, p. 10.
2. Miller, W. J. and W. H. Stone. 1959. Desorption- A technique which increases the yield of reagents from absorptions. The Immunogenetics Letter Vol. 1, No. 2, p. 10.
3. Miller, W. J. 1961. Presence of species-specific antigens in tissues. The Immunogenetics Letter Vol. 2, No. 5, p. 1012.
4. Miller, W. J. and B. G. Morris. 1961. A fertile heifer with blood-type mosaicism co-twin with a bull. The Immunogenetics Letter Vol. 2, No. 6, p. 44.
5. Grundbacher, F. J. and W. J. Miller. 1961. Production of anti-cattle U reagents in rabbits. The Immunogenetics Letter Vol. 2, No. 6, pp. 49-50.
6. Miller, W. J. and C. Stormont. 1962. Probability of the first versus the second bull being the sire in parentage cases. The Immunogenetics Letter Vol. 2, No. 7, pp. 78-79.
7. Braend, M., W. J. Miller and C. Stormont. 1962. The new reagent for the M blood group system of cattle. The Immunogenetics Letter Vol. 2: No. 7, pp. 78-79.
8. Miller, W. J., D. R. Osterhoff and C. Stormont. 1962. The two new reagents for the B blood group system of cattle. The Immunogenetics Letter 2: pp. 136-138.
9. Miller, W. J. and M. L. Wendorf. 1962. Antigenic differences of the erythrocytes among ring neck pheasants. The Immunogenetics Letter Vol. 2, No. 7, pp. 93-95.
10. Miller, W. J. 1963. Individual differences of the red cells of ring neck doves detected by seed extracts. The Immunogenetics Letter Vol. 3, No. 2, pp. 57-61.
11. Asmundson, V. S., W. J. Miller and F. X. Ogasawara. 1963. Tests for interaction specificities of species crosses in poultry. The Immunogenetics Letter Vol. 3, No. 1, pp. 29-45.
12. Miller, W. J. 1964. Are lectin detected "antigens" serological antigens?. The Immunogenetics Letter Vol. 3, No. 4, pp. 181-184.
13. Miller, W. J. 1964. First linkage of a species antigen in the genus Streptopelia. The Immunogenetics Letter Vol. 3, No. 3, pp. 105-111.
14. Gregory, P. W. and W. J. Miller. 1965. Coat color, pattern, horn condition, and blood groups in cattle. The Immunogenetics Letter Vol. 4, No. 1, pp. 15-19.
15. Miller, W. J. 1966. Frequencies of blood groups in American Ayrshire and Brown Swiss cattle. The Immunogenetics Letter Vol. 4, No. 4, p. 192-195.
16. Hollander, W. F. and W. J. Miller. 1982. A new sex-linked mutation, "Web-lethal". American Racing Pigeon News Oct. 1982 pp. 50-51.
17. Franklin, W. J., P. J. Hollander, and W. J. Miller. 1984. Blood typing studies of Llamas and Guanacos. International Llama Association Newsletter.
18. Miller, W. J. 1998. Miller returns from Brazil. The Doveline July/August p. 7.
19. Hollander, W. F. and W. J. Miller. 1999. Where do new varieties of ringneck doves come from?. The Doveline Jan./Feb. p. 3.

Speeches-

1. Miller, W. J. 1978 Blood groups in domestic animals. III Semana de Zootecnia Palestra: GRUPOS SANGUINEOS NOS ANIMAIS DOMESTICOS. no Curso de Melhoramento Animal, USP Faculdade de Medicina Veterinaria e Zootecnia, Centro de Zootecnia e Industrias Pecuarias "Fernando Costa" , Pirassununga , S.P., BRASIL
2. Miller, W. J. 1984. Bird Genetics: Generalizations and specifics in cage birds. 18 Oct. ‘84. Invited talk for the Mid America Cage Bird Society, Des Moines, IA.
3. Miller, W. J. 1985. Blood typing and its role in breed history evaluation. 29 Sept. ‘84. Invited talk for the annual meeting and conference American Minor Breeds Conservancy, Sedgwick County Zoological Society, Wichita, Kansas.
4. Miller, W. J. 1985. Seminar on Dairy Goat Genetics at the annual meeting of the Iowa Dairy Goat Association. 26 Oct. ‘85. School of Veterinary Medicine, Ames, IA.
5. Miller, W. J., D.A.A. Oliveira, L. E. L. Pinheiro and M. Y. Kuabara 1993 Potential application of blood groups to production characters of cattle. X Congresso Brasileiro de Reprodução Animal, Anais Vol II:117-124. Belo Horizonte, M. G., BRASIL
6. Miller, W.J. 1996 . Some neglected advances in classical genetics. I Simpósio Internacional de Monogástricos. Universidade Federal Rural do Rio de Janeiro, Itaguaí, Rio de Janeiro, Brasil, Sept. , 16 pp.
7. Miller, W. J. 1997. Cage Birds in Brazil. 26 Oct. ‘97. Invited talk for the Mid-America Cage Bird Society at The Botanical Center, Des Moines, IA. 

Letter to the Editor-

1. Miller, W. J. 1950. Maintaining species purity in nature. North American Veterinarian 31: 109.

 Grant Reviews-

1. Reviewed Grant Proposal Section 406, PL 89-106 808 University of Florida, Gainsville on Development of a subunit diagnostic test for bovine Babesiosis.

 Committees-

Departmental:

• 1973-1978: Safety Inspection- United Way
• 1973-1977: Immunobiology Committee
• 1975-1987(?): Staff Meeting Secretary
• 1976-1987(?): Graduate Student Teaching Assignments...

 University:

• 1976: Graduate Faculty Membership
• 1976-1977: Laboratory Animal Facilities Committee

 Outside Organizations:

• 1978: Editorial Committee on the Pigeon Science and Genetics Newsletter

 Research Projects-

• 1974-78: Genetic, Developmental, and Pathological Aspects of Spinal Dysraphism in the Dog. Co-operator with Drs. Draper and Kluge in Vet. Med. 43013-11-23-0002, $5,678 Immunogenetics- 102-4051-51-1940
• 1979-80 Faculty Improvement Leave. Immunogenetics 102-4051-51-2251 continued at Universidade Estadual de São Paulo, Jaboticabal. A cattle blood typing laboratory was initiated there.
• 1980-82 Iowa Agriculture and Home Economics Experiment Station, Project No. 102-4051-51-2251, $4,000. Immunogenetics of Vertebrates.
• 1982 The International Llama Association made a $2,000 mini-research grant available to Wilmer J. Miller, Paul Hollander and Bill Franklin.

Awards-

• 1978: Received the "W. F. Hollander Merit Award" by C. F. Graefe for pigeon genetics research.

Graduate Students-

1964 Clarence Goon, M.S.
1968 Derald E. Kimm, M.S.
1973-78 Carol Stieler Halverson, Ph.D.
1976 Bin Kikuchi
1979 Karen Collias, M.S.

Undergraduate Students-

Domingo Cruz Filho
Luzia Helena Queiroz
Claudemir de Carvalho

Graduates-
Ana Isabel de Assis
Miriam R. Luz Gianonni

1980-81 Undergraduates-
3 Biology Majors

Graduates-
Irma Carillo, Genetics
1982-86 Undergraduates: 4-10

External Examiner-

July 1984 Served as external examiner for Miss Marion Wanjiku Wangai’s M.S. thesis on blood groups in camels, University of Nairobi, Kenya.

Courses Taught at Iowa State-

Genetics 301- Introductory Genetics
Genetics 320- Introductory Genetics
Genetics 340- Principles of Genetics
Genetics 350- Principles of Genetics
Genetics 546- Immunogenetics
Genetics 646- Immunogenetics
Genetics 260- Heredity and Society
Genetics 360- Heredity and Society
Genetics 490- Independent Study
Genetics 699- Research
Genetics 690- Seminar
Biology 490
IMBIO 690
IMBIO 699

1963 Fall- GEN 301 3 Cr. 25 students
Winter- GEN 301 3 Cr. 77 students
1964 Spring- GEN 301 3 Cr. 69 students
Fall- GEN 646 3 Cr. 5 students
Fall- GEN 301 3 Cr. 128 students
1965 Fall- GEN 646 3 Cr. 5 students
Winter- GEN 301 3 Cr. 110 students
Spring- GEN 301 3 Cr. 101 students
1966 Winter- GEN 301 3 Cr. 82 students
1967 Fall- GEN 646 3 Cr. 4 students
Winter- GEN 301 3 Cr. 97 students
Spring- GEN 301 3 Cr. 110 students
1968 Fall- GEN 301 3 Cr. 107 students
Winter- GEN 301 3 Cr. 97 students
Spring- GEN 301 3 Cr. 111 students
1969 Fall- GEN 646 3 Cr. 9 students
Winter- GEN 301 3 Cr. 92 students (Sec. 1)
Winter- GEN 301 3 Cr. 102 students (Sec. 2)
1970 Winter- GEN 301 3 Cr. 107 students
Spring- GEN 301 3 Cr. 141 students (Sec. 2)
Spring- GEN 301 3 Cr. 138 students (Sec. 3)
1971 Fall- GEN 646 5 Cr. 5 students
Winter- GEN 301 3 Cr. 110 students
Spring- GEN 301 3 Cr. 109 students
1972 Winter- GEN 301 3 Cr. 102 students
1973-74 Fall- GEN 546 3 Cr. 9 students
Winter- GEN 340 3 Cr. 75 students
Spring- GEN 350 3 Cr. 140 students
1975-76 Fall- GEN 340 3 Cr. 93 students
Fall- GEN 546 3 Cr. 18 students
Winter- GEN 340 3 Cr. 222 students
Spring- GEN 360 3 Cr. 18 students
1976-77 Fall- GEN 546 3 Cr. 14 students
Winter- GEN 340 3 Cr. 136 students
Spring- GEN 340 3 Cr. 136 students
Winter- GEN 490 3 Cr. 1 student (Independent study)
Winter- GEN 699 3 Cr. 2 graduate students
1977-78 Summer- BIO 490 3 Cr. 1 student
Fall- IMBIO 699 1 Cr. 1 student
Fall- GEN 546 3 Cr. 5 students
Winter- GEN 340 3 Cr. 156 students
Winter- GEN 490 4 Cr. 1 student
Winter- IMBIO 600 1 Cr. 1 student
Spring- GEN 340 3 Cr. 134 students
Spring IMBIO 699 1 Cr. 1 student
1978-89 Summer- GEN 340 3 Cr. 37 students
1980-81 Summer GEN 340 3 Cr. 44 students
Fall- GEN 599 3 Cr. 1 student
Fall- GEN 690 1 Cr. 12 students
Winter- GEN 340 3 Cr. 104 students
Winter- GEN 599 5 Cr. 1 student
Spring- GEN 340 3 Cr. 176 students
Spring- GEN 599 3 Cr. 1 student
1982-83 Summer- GEN 320 3 Cr. 48 students
Spring- GEN 320 3 Cr. 129 students
Fall- GEN 320 3 Cr. 89 students

Summer- GEN 490 Cr. 1 student
Fall- GEN 490 3 Cr. 2 students
1983-84 Spring- GEN 320 3 Cr. 115 students
Summer- GEN 320 3 Cr. 43 students
Summer- GEN 490 3 Cr. 2 students
Summer- GEN 490 6 Cr. 1 student
Fall- GEN 320 3 Cr. 91 students
1984-85 Spring- GEN 320 3 Cr. 100 students
Summer- GEN 490 3 Cr. 2 students
Summer- GEN 320 3 Cr. 32 students
Fall- GEN 590 3 Cr. 8 students
Fall- GEN 690 1 Cr. 12 students
Spring- GEN 320 3 Cr. 94 students
Summer- GEN 320 3 Cr. 27 students
1985-86 Fall- GEN 320 3 Cr. 83 students
Fall- GEN 490 3 Cr. 1 student
Spring- GEN 320 3 Cr. 80 students
Summer- GEN 320 3 Cr. 36 students
1986-87 Fall- GEN 260 3 Cr. 31 students
Spring- GEN 320 3 Cr. 116 students
Summer- GEN 320 3 Cr. 30 students
Fall- GEN 320 3 Cr. 123 students
N.B: Additional students to 1994 are not added.

Supervised Visiting Professor for 14 months, Dr. Indalecio Rodolfo Quinteros, Professor Titular Full-time, Catedra Genetica y Biometrica, Director del Instituto de Immunogenetica Animal y Genetica. 

During his teaching years at ISU, Dr. Miller has essentially completed a survey of Introductory Genetics for college students in the upper class (Junior-Senior) level. This is aimed at the ISU course Genetics 320. These genetics notes contain 291 pages. The primary purpose is to obviate the need for furious scribbling by students in taking lecture notes, allowing them to pay attention. The secondary purpose is to correct the misleading aspects of the manipulation of Mendelism in textbooks. See the following list of improvements: 

How Dr. Miller’s text "A Survey of Genetics" improves on other Genetic textbooks: Other texts

1. Much more consistent in appropriate gene symbol usage. Very poor

2. Distinguishes dominance from epitasis, and alleles from non-alleles on a classical basis. Do not

3. Note the 3 kinds of epitasis. Do not

4. More arithmetic methodology leading to much more rapid than the checkerboard with Mendeliam problems. Some, but limited solutions

5. Handles mixed mating types (not just F2 or testcrosses). No examples

6. Includes Pedigree Analysis with genetic interactions and with generalizations and algorithmic method given. None

7. More complete and generalized chapter on blood types. Brief or missing

8. Single, comprehensive pedigree (network) style adaptable to all uses and fitting the blackboard better. (Human style square and circles also given.) This style limited to calculations of inbreeding and relationships.

9. Half of the sample questions and models are original and new, adapted from published research reports and, in a few cases, the author's own research.

Dr. Miller has developed 800+ multiple choice questions on computer file suitable for test scoring in Genetics 320. These notes have since turned into a text book written by Dr. Miller and published by Ginn press, 328 pp.